Typification in and contributions to a revision of Psammisia (Ericaceae: Vaccinieae)


  • James L. Luteyn The New York Botanical Garden




Since the last complete revision of Psammisia (Ericaceae: Vaccinieae) in 1932 many new species, synonyms, and local treatments have been published and genetic information is slowly becoming available. This paper is based on field and herbarium observations made over a period of nearly 50 years and helps to clarify questions of typification and species relationships towards an updated revision. Emphasis is herein given to species occurring in Peru and Bolivia for which a key to the species occurring in that region is made available for the first time. Notes on the poorly understood yet historically important collections of Ruiz & Pavón, Humboldt & Bonpland, Mathews, the Schomburgk brothers, and Linden, Funck, & Schlim are provided as they refer to South American Ericaceae. Neotypes for Thibaudia formosa and Thibaudia nutans are also included.


Alexander, S. 2011. Making sense of the Schomburgk expeditions. The Plant Press 14(2), April 2011.

Ballington, J.R., J.L. Luteyn, M.M. Thompson, K. Romoleroux, & R. Castillo. 1993. Rubus and vacciniaceous germplasm resources in the Andes of Ecuador. Pl. Genetic Res. Newslett. 93:9–15.

Barreiro, A.J. 1931. Relación del viaje hecho á los reynos de Perú y Chile pos los botánicos y dibuxantes enviados por el Rey para aquella expedición, extractada de los diarios por el orden que llevo a cabo su autor dn. Hipólito Ruiz. [First Spanish transcription of the oldest known version of H. Ruiz’ original unpublished field journals, revised and annotated with an epilogue and official documents by A.J. Barreiro.]

Bentham, G. 1838. Enumeration of the plants collected by Mr. Schomburgk, British Guiana. Ann. Nat. Hist. 2(8):105–111. [explanation of original Rob. Schomburgk numbering series]

Bentham, G. 1839–57. Plantae hartwegianae. London. [Ericaceae, 1846:219–226]

Bentham, G. & J.D. Hooker. 1876. Genera plantarum: ad exemplaria imprimis in Herberiis Kewensibus servata definita. A. Black, London. Https://doi.org/10.5962/bhl.title.747

Brako, L. & J.L. Zarucchi. 1993. Catalogue of the flowering plants and gymnosperms of Peru. (Catálogo de las angiospermas y gimnospermas del Perú). Monogr. Syst. Bot. Missouri Bot. Gard. 45:1–1286.

Britton, N.L. 1921. Studies of West Indian plants –X. Bull. Torrey Bot. Club 48:327–343. [Psammisia recurvata Britton, p. 335]

Brummitt, R.K. & C.E. Powell, eds. 1996. Authors of plant names. Kew Publishing, London.

Burdet, H.M. 1979. Auxilium ad botanicorum graphicem. Conservatoire et Jardin botaniques de la Ville de Genève.

Candolle, A.P. de. 1823–1873. Prodromus systematis naturalis regni vegetabilis, sive enumeratio contracta ordinum, generum specierumque plantarum hucusque cognitarum, juxta methodi naturalis normas digesta. 17 vols. Treuttel et Würtz, Paris, Strasbourg & London.

Chaudhri, M.N., I.H. Vegter, & C.M. dewal. 1972. Index herbariorum: part II(3), Collectors, I–L. Regnum Veg. 86. A. Oosthoek’s Uitgeversmaatschappij N.V., Utrecht.

Cronquist, A. 1978. Once again, what is a species? Pp. 3–20. In: J.A. Romberger, ed. Biosystematics in Agriculture. Allanheld, Osmun & Co., Montclair, New Jersey.

Dahlgren, B.E. 1940. Travels of Ruiz, Pavón, and Dombey in Peru and Chile (1777–1788). Field Mus. Nat. Hist., Bot. Ser. 21. Publ. 467. [English translation of Barreiro (1931)] https://doi.org/10.5962/bhl.title.2305

Decaisne, J. 1854a. Rev. Hort., sér. IV, Tome 3(1):5–6, fig. 1 (1 Jan 1854). [Psammisia penduliflora (Dunal) Klotzsch]

Decaisne, J. 1854b. Rev. Hort., sér. IV, Tome 3(10):181, fig. 10 (15 May 1854). [Psammisia sarcantha Decne.]

Decaisne, J. 1854c. Rev. Hort., sér. IV, Tome 3(10):182 (15 May 1854). [“Psammisia, Planchoniana Decne.”]

Don, G. 1831–1838. General history of the dichlamydeous plants. 4 vols. J.G. and F. Rivington et al., London. [Ericaceae Tribe III. Vaccinieae, vol. 3:851–863. 1834] https://doi.org/10.5962/bhl.title.16519

Dunal, M.F. 1839. Vaccinieae. In: A.P de Candolle, ed. Prodromus systematis naturalis regni vegetabilis, 7(2):552–579. Treuttel & Würtz, Paris.

Fedtschenko, B.A. & N.A. Basilevskaja. 1926. Ericaceae austroamericanae novae. Bot. Mater. Gerb. Glavn. Bot. Sada SSSR 6:21–26. [p. 24, Macleania crenulata B.Fedtsch. & Basil.]

Foster, R.C. 1958. A catalogue of the ferns and flowering plants of Bolivia. Contr. Gray Herb. 184:1–223.

Grimé, W.E. & T. Plowman. 1987. Type photographs at the Field Museum of Natural History. Taxon 36:425–426. [URL: http://www.jstor.org/stable/1221436]

Guillemin, J.A. 1833. Archives de botanique; ou, Recueil mensuel de mémoires originaux, d'extraits et analyses bibliographiques, d'annonces et d'avis divers concernant cette science; rédigées par une société de botanistes français et étrangers, sous la direction de m. A.-J. Guillemin. 2 vols. Bureau des archives, Paris. [vol. 1, pl. Facing p. 380—Ruiz’ signature]

Hiepko, P. 2006. Humboldt, his botanical mentor Willdenow, and the fate of the collections of Humboldt & Bonpland. Bot. Jahrb. Syst. 126:509–516.

Hoerold, R. 1909a. Systematische Gliederung und geographische Verbreitung der amerikanischen Thibaudieen. Bot. Jahrb. Syst. 42:251–334.

Hoerold, R. 1909b. Ericaceae. Pp. 92–94. In: E. Ule, III. Beitrage zur Flora der Hylaea nach den Sammlungen von Ule’s Amazonas-Expedition. Verh. Bot. Vereins Prov. Brandenburg 50:69–124. [Psammisia ulei Hoerold, pp. 92–93]

Hooker, W.J. 1837. Icon. Pl. 2: tab. 109. [Macleania Hook.]

Hooker, W.J. 1839. Icon. Pl. 3: tab. 243. [Anthopterus Hook.]

Hooker, W.J. 1847. Bot. Mag. 73: tab. 4344. [Thibadia pichinchensis Benth. Var. Glaber Hook.]

Hooker, W.J. 1860. Bot. Mag. 86: tab. 5204. [Psammisia penduliflora Hook.]

Hooker, W.J. 1864. Bot. Mag. 90: tab. 5450. [Thibaudia sarcantha Hook.]

Hooker, J.D. 1865. Bot. Mag. 91: tab. 5526. [Psammisia longicolla Hook.f.]

Hooker, J.D. 1865. Bot. Mag. 91: tab. 5547. [Thibaudia jessicae Hook.f.]

Hooker, J.D. 1876. Vacciniaceae. In: Bentham, G. & J.D. Hooker. Genera plantarum 2:564–577. London.

Hooker, J.D. 1878. Bot. Mag. 104: tab. 6393. [Eurygania ovata Hook.f.]

Index Herbariorum. 2016. A global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium [http://sweetgum.nybg.org/ih/].

Jaramillo-Arango, J. 1952. Relación del viaje hecho á los reynos de Perú y Chile por los botánicos y dibuxantes enviados por el Rey para aquella expedición, extractada de los diarios por el orden que llevo a cabo su autor dn. Hipólito Ruiz. Edition 2. Madrid. [Second Spanish edition of H. Ruiz’ field journals transcribed by J. Jaramillo-Arango, this time incorporating Ruiz’ unpublished second and third versions of the original, respectively, “Compendio del viage que hizo á los reynos del Perú y Chile el botánico Español dn. Hipólito Ruiz en el año de 1777 hasta el de 1788” and “Relación histórica del viage, que hizo á los reynos del Perú y Chile el botánico dn. Hipólito Ruiz en al año de 1777 hasta el de 1788, en cuya época regresó a Madrid.”]

JSTOR-Plants. 2018. JSTOR global plants. Http://plants.jstor.org/.

Killip, E.P. 1927. Report of the Killip-Smith botanical expedition to Colombia, 1926–1927. J. New York Bot. Gard. 28(333):205–220. [see J.H. Barnhart’s footnote 3, p. 210–212, for notes on Linden, Funck, & Schlim]

Klotzsch, J.F. 1848[1849]. Ericaceae. Pp. 1087–1088. In: Rich. Schomburgk, ed. Reisen in Britisch-Guiana in den Jahren 1840–1844: nebst einer Fauna und Flora Guiana's nach Vorlagen von Johannes Müller, Ehrenberg, Erichson, Klotzsch, Troschel, Cabanis und Andern, 3 vols. J.J. Weber, Leipzig. [Ericaceae, vol. 3, actual 7–10 Mar 1849]

Klotzsch, J.F. 1851. Studien über die natürliche Klasse Bicornes Linné. Linnaea 24:1–88.

Kron, K.A., E.A. Powell, & J.L. Luteyn. 2000 [Abstract]. Molecular systematics of Macleania and Psammisia and their relationship to other members of Vaccinieae. Amer. J. Bot. 87(Supplement):137.

Kron, K.A., E.A. Powell, & J.L. Luteyn. 2002a. Phylogenetic relationships within the blueberry tribe (Vaccinieae, Ericaceae) based on sequence data from matk and nuclear ribosomal ITS regions, with comments on the placement of Satyria. Amer. J. Bot. 89:327–336. Doi: 10.3732/ajb.89.2.327

Kron, K.A., W.S. Judd, P.F. Stevens, D.M. Crayn, A.A. Anderberg, P.A. Gadek, C.J. Quinn, & J.L. Luteyn. 2002b. Phylogenetic classification of Ericaceae: molecular and morphological evidence. Bot. Rev. 68:335–423.

Kunth, C.S. 1819. Ericaceae. Pp. 263–299, plates 248–264. In: F.H.A. von Humboldt, A.J. Bonpland, & C.S. Kunth. 1815–1825. Nova genera et species plantarum. 3 vols. Lutetiae Parisiorum [Paris].[“Ericeae,” vol. 3, Fasc. 11:263–288 (Jul 1819); Fasc. 12:289–299 (Nov 1819); plates 248–264 (Jul 1819)]

Lack, H.W. 2004. The botanical field notes prepared by Humboldt and Bonpland in tropical America. Taxon 53:501–510. Http://www.jstor.org/stable/4135629

Lechler, W. 1857. Berberides Americae australis. E. Schweizerbart, Stuttgartiae. 59 pp. [incl. Lechler’s exicattae on pp. 49–59, with A.H.R. Grisebach’s Ericaceae nom. Nud. On p. 58]

León, B. 2006. Ericaceae endémicas del Perú. In: B. León et al., eds. El libro rojo de las plantas endémicas del Perú. Revta. Peruana Biol., Número especial 13(2):285s–293s. Facultad de Ciencias Biológicas UNMSM.

Levin, D.A. 1979. The nature of plant species. Science n.s. 204(4391):381–384.

Linden, J. 1851. Établissement de botanique et d’horticulture de J. Linden, a Luxembourg. Prix-courant pour printempts 1847. M. Behrens fils, Luxembourg. (TL-2 abbrev.: Établ. Linden, prix-courant)(loosely translated: Prix-courant de l'établissement d'introduction pour les plantes nouvelles; and often referred to simply as Linden’s Catalogue, ex. Lind. Cat. No. 6:4.1851). [Thibaudia schlimiana Linden, 6:4. 1851]

Linden, J. & J.E. Planchon. 1863. Plantae columbianae. Bruxelles. Reprint ed. 1977, Boerhaave Press, Leiden.

Lindley, J. 1836. Bot. Reg. 21: sub pl. 1791. [Cavendishia nobilis Lindl.]

Luteyn, J.L. 1981. Notes on Neotropical Vaccinieae (Ericaceae), IX. Seven new Andean blueberries. Brittonia 33:371–381.

Luteyn, J.L. 1983. Part I. Cavendishia. Fl. Neotrop. Monogr. 35:1–290. [Cavendishia nobilis Lindl., pp. 2–4]

Luteyn, J.L. 1987. New species and notes on Neotropical Ericaceae. Opera Bot. 92:109–130.

Luteyn, J.L. 1991. Key to the subfamilies and genera of Neotropical Ericaceae. Nordic J. Bot. 11:623–627.

Luteyn, J.L. 1996. 147. Ericaceae. In: G. Harling & L. Andersson, eds. Flora of Ecuador 54:1–404, color plates I–VIII.

Luteyn, J.L. 1998. Ericaceae. In: J.A. Steyermark et al., eds. Flora of the Venezuelan Guayana 4:735–769. Missouri Botanical Garden.

Luteyn, J.L. 2002a. Diversity, adaptation, and endemism in Neotropical Ericaceae: biogeographical patterns in the Vaccinieae. In: K. Young, C. Ulloa Ulloa, J.L. Luteyn & S. Knapp, coauthors & eds. Plant evolution and endemism in Andean South America. Bot. Rev. 68:55–87.

Luteyn, J.L. 2002b. Key to the species of Ericaceae of Bolivia, including two new species. Sida 20:1–20.

Luteyn, J.L. 2007. Ericaceae of Ecuador. Rapid Color Guide # 203 versio?n 1. 12/2007. The Field Museum. [www.fmnh.org/plantguides/]

Luteyn, J.L. 2018. Towards a taxonomic treatment of the Ericaceae for Bolivia and Peru: new species of Thibaudia, synonyms, lectotypes, and ob servations. J. Bot. Res. Inst. Texas 12:89–114.

Luteyn, J.L. 2019. In prep. Monograph of the neotropical genus Macleania (Ericaceae: Vaccinioideae: Vaccinieae).

Luteyn, J.L. & C. Maldonado. 2014. Ericaceae. In: P.M. Jørgensen et al., eds. Catálogo de las plantas vasculares de Bolivia. Monogr. Syst. Bot. Missouri Bot. Gard. 127:576–581.

Luteyn, J.L. & P. Pedraza-Peñalosa. 2007a. Neotropical blueberries: The plant family Ericaceae.[www.nybg.org/bsci/res/lut2]. Superseded by Luteyn & Pedraza-Peñalosa 2012–onward.

Luteyn, J.L. & P. Pedraza-Peñalosa. 2007b. Santuario histórico de Machu Picchu, Cusco, Perú: Ericaceae de Machu Picchu. Rapid Color Guide # 238 versio?n 1. 12/2007. The Field Museum. [http://fm2.fieldmuseum.org/plantguides/guideimages.asp?ID=348]

Luteyn, J.L. & P. Pedraza-Peñalosa. 2007c. Ericaceae de Bolivia. Rapid Color Guide # 239 versio?n 1. 12/2007. The Field Museum. [http://fm2.fieldmuseum.org/plantguides/guideimages.asp?ID=349]

Luteyn, J.L. & P. Pedraza-Peñalosa. 2012-onward. Blueberry relatives of the New World tropics (Ericaceae) (http://sweetgum.nybg.org/ericaceae/index.php). The New York Botanical Garden, Bronx, New York. [this version not cleared by JLL, nor later versions by PP-P]

Luteyn, J.L. & P. Pedraza-Peñalosa. In prep. Ericaceae of Bolivia and adjacent southern Peru.

Luteyn, J.L. & D.S. Sylva S. 1999. “Murri" (Antioquia Department, Colombia): hotspot for Neotropical blueberries (Ericaceae: Vaccinieae). Brittonia 51:280–302.

Luteyn, J.L. & M.L. Vidal-Lemus. 2015. Ericaceae. In: J.L. Fernández Alonso & J. Aguirre Ceballos, eds. La flora de la Real Expedición Botánica del Nuevo Reyno de Granada (1783–1816), Tome XXI [Flora de Mutis]:1–36, lam. I–XLIII. Instituto Colombiano de Antropología e Historia (ICANH). Bogotá, República de Colombia.

Luteyn, J.L. & R.L. Wilbur. 2005. Ericaceae. In: Wm. Burger, ed. Flora Costaricensis. Fieldiana, Bot., n.s. 45:i–vi, 1–107. Publ. 1536.

Luteyn, J.L. & R.L. Wilbur. 2010. Ericaceae. Pp. 239–283. In: B.E. Hammel et al., eds. Manual de las plantas de Costa Rica. Missouri Botanical Garden.

Luteyn, J.L., E.M. Ortiz, & B. León. 2008a. Notes on and lecotytpification of Augusto Weberbauer’s collections of Peruvian Ericaceae. Revta. Peruana Biol. 15:127–134.

Luteyn, J.L., R.L. Wilbur, & L.J. Dorr. 2008b. Ericaceae. In: G. Davidse et al., eds. Flora Mesoamericana 4(1):411–463. Universidad Nacional Autónoma de Mexico, Missouri Botanical Garden, and The Natural History Museum (London).

Macbride, J.F. 1944. Vaccinium and relatives in the Andes of Peru. Univ. Wyoming Publ. Sci.11:37–46.

Macbride, J.F. 1959. Ericaceae. In: J.F. Macbride & B.E. Dalhgren, eds. Flora of Peru. Field Mus. Nat. Hist., Bot. Ser. 13(V,1):50–149.

Maguire, B., J.A. Steyermark, & J.L. Luteyn. 1978. Ericaceae. In: B. Maguire et collab., eds. Botany of the Guayana Highland. Mem. New York Bot. Gard. 29:139–203.

Mansfeld, R. 1925. Beiträge zur Kenntnis der südamerikanischen Ericaceen. I. Notizbl. Bot. Gart. Berlin-Dahlem 9:435–442. Doi: 10.2307/3994409

Mcneill, J. 2014. Holotype specimens and type citations: general issues. Taxon 63:1112–1113. Http://dx.doi.org/10.12705/635.7

Meissner, C.F. 1840–1906. In: K.F.P. von Martius, Flora Brasiliensis, enumeratio plantarum in Brasilia hactenus detectarum: quas suis aliorumque botanicorum studiis descriptas et methodo naturali digestas partim icone illustratas. R. Oldenbourg, Monachii et Lipsiae [Munich & Leipzig]. [Ericaceae s.l. Vol. 7:127–172 (publ. 10 July 1863)]

Miller, H. 1970. The herbarium of Aylmer Bourke Lambert: notes on its acquisition, dispersal, and present whereabouts. Taxon 19:489–553. Http://www.jstor.org/stable/1218947

Morton, C.V. 1971. The fern collections in some European herbaria, VII: Brussels. Amer. Fern J. 61:59–75. [see pp. 72–73 for notes on Linden, Funck, & Schlim]

Nevling, L.I., Jr. 1970. The ecology of an elfin forest in Puerto Rico, 12. A new species of Gonoccalyx (Ericaceae). J. Arnold Arbor. 51:221–227. [see pp. 224–227 for notes on Linden, Funck, & Schlim]

Niedenzu, F. 1890. Über den anatomischen Bau der Laubblätter der Arbutoideae und Vaccinioideae in Beziehung zu ihrer systematischen Gruppierung und geographischen Verbreitung. Bot. Jahrb. Syst. 11:134–263. [“Ceratostema mutans [sic]” nom. Nud., pp. 211 and 224; “Th. Guianensis Kl.,” pp. 208 and 224]

Pedraza-Peñalosa, P. 2015a. Ericaceae. Pp. 448–460. In: P.A. Idárraga et al, eds. Flora de Antioquia: Catálogo de las plantas vasculares. Vol. II. Listado de las plantas vasculares del Departamento de Antioquia. Programa Expedición Antioquia-2013. Serie Biodiversidad y Recursos Naturales. Universidad de Antoquia, Missouri Botanical Garden & Oficina de Planeación Departamental de la Gobernación de Antioquia. D’Vinni and Bogotá.

Pedraza-Peñalosa, P. 2015b. New blueberry and mortiño relatives (Ericaceae) from northwestern Colombia. Phytokeys 49:33–58. Doi: 10.3897/phytokeys.49.8383

Pedraza-Peñalosa, P. 2015c[2016]. Three new species of Psammisia s.l. (Ericaceae: Vaccinieae), blueberry relatives endemic to Colombia. Brittonia 68:25–36 [Published online: 11 November 2015]. Doi: 10.1007/s12228-015-9388

Pedraza-Peñalosa, P., N.R. Salinas, & W.C. Wheeler. 2013. Venation patterns of neotropical blueberries (Vaccinieae: Ericaceae) and their phylogenetic utility. Phytotaxa 96:1–53. Doi: 10.11646/phytotaxa.96.1.1

Pedraza-Peñalosa, P., N.R. Salinas, A.L.S. Virnig, & W.C. Wheeler. 2015. Preliminary phylogenetic analysis of the Andean clade and the placement of new Colombian blueberries (Ericaceae, Vaccinieae). Phytokeys 49:13–31. Doi: 10.3897/phytokeys.49.8622

Planchon, J.E. 1853. Flore des serres, et des jardins de l’Europe; on descriptions des plantes les plus rares et les plur méritantes, … et extraits des Botanical Magazine, Botanical Register, Paxton’s Magazine of Botany, etc. Ghent 8:205, pl. 825. [Psammisia sclerophylla Planch. & Linden]

Powell, E.A. & K.A. Kron. 2003. Molecular systematics of the northern Andean blueberries (Vaccinieae, Vaccinioideae, Ericaceae). Intern. J. Pl. Sci. 164:987–995.

Regel, ed. 1874. J. Linden und dessen Etablissement in Gent. Gartenflora 23:196–201.

Restrepo P., D.L., D.S. Sylva S., G.E. Martínez A. & C.E. Orrego P. 1989. Estudio florístico de la cuenca alta del Río Cuevas, Municipio de Frontino, Antioquia. Unpubl. Manuscript. Department of Biology, Universidad Nacional de Colombia, Seccional Medellín.

Ruiz, H. & J. Pavón. 1794. Flora peruvianae, et chilensis prodromus, sive novorum generum plantarum peruvianarum et chilensium descriptiones, et icones. Descripciones y láminas de los nuevos géneros de plantas de la flora del Perú y Chile. Impr. De Sancha, Madrid.

Ruiz, H. & J. Pavón. 1798. Systema vegetabilium florae peruvianae et chilensis, characteres prodromi genericos differentiales, specierum omnium differentias, durationem, loca natalia, tempus florendi, nomina vernacula, vires et usus nonnullis illustrationibus interspersis complectens. Typis Gabrielis de Sancha, Madrid.

Ruiz, H. & J. Pavón. 1798–1802[1956–1960]. Flora peruviana et chilensis, sive descripciones, et icones, plantarum peruvianarum, et chilensium, secundeum systema linnaeanum digestae, cum characteribus plurium generum evulgatorum reformatis, 3 vol. Typis Gabrielis de Sancha, Madrid. [Volumes 4 and 5 were published in the Anales Inst. Bot. Cavanilles 13–17. 1956–1960; Ericaceae text in 14:756–767(1957) and plates 383–388 in 14:779–784(1957); Thibaudia coarctata text 14:762 and plate 385 14:781]

Rusby, H.H. 1912. New species from Bolivia, collected by R.S. Williams—2. Bull. New York Bot. Gard. 8(No. 28 November 23, 1912):89–136 [“Issued separately, in advance, 19 N 1912”]. [Ericaceae/Vaccinieaceae, pp. 110–111]

Saint-Hilaire, J. 1805. Exposition des familles naturelles et de la germination des plantes, par Jaume Saint-Hilaire; contentant la description de deux mille trois cent trente-sept genres, et d'environ quatre mille espèces, les plus utiles et les plus interessantes. Cent dix-sept planches dont les figures ont été dessinées par l’auteur, représentent les caractères des familles et les differens modes de germination des plantes monocotylédones et dicotlédones. 2 vols. Treuttel et Würtz, Paris. [“Les Bruyeres” incl. Vaccinium, Thibaudia, and Ceratostema are found in vol. 1:362–363]

Sayre, G. 1975. Cryptogamae exsiccatae an annotated bibliography of exsiccatae of algae, lichenes, hepaticae, and musci, V. Unpublished exsiccatae. I. Collectors. Mem. New York Bot. Gard. 19(3):277–423.

Schomburgk, R., ed. 1847–1848[1849]. Reisen in Britisch-Guiana in den Jahren 1840–1844: nebst einer Fauna und Flora Guiana's nach Vorlagen von Johannes Müller, Ehrenberg, Erichson, Klotzsch, Troschel, Cabanis und Andern, 3 vols. J.J. Weber, Leipzig. [Vol. 3 (actual 7–10 Mar 1849) was entitled "Versuch einer Fauna und Flora von Britisch-Guiana,” basically a synopsis of the flora of British Guiana.] Https://doi.org/10.5962/bhl.title.109982.

Schultes, R.E. & M.J. Nemry von Thenen de Jaramillo-Arango. 1998. The journals of Hipólito Ruiz: Spanish botanist in Peru and Chile 1777–1788. Timber Press, Portland, Oregon. [English translation of Jaramillo-Arango (1952)]

Sleumer, H.O. 1934. Ericaceae americanae novae vel minus cognitae, I. Notizbl. Bot. Gart. Berlin-Dahlem 12:119–140. [Thibaudia sanmartensis Sleum.]

Sleumer, H.O. 1935. Ericaceae americanae novae vel minus cognitae II. Notizbl. Bot. Gart. Berlin-Dahlem 12:277–294.

Sleumer, H.O. 1936. Die Arten der Gattung Vaccinium L. In Zentral- und Südamerika. Notizbl. Bot. Gart. Berlin-Dahlem 13:111–140.

Sleumer, H.O. 1941. Vaccinioideen-Studien. Bot. Jahrb. Syst. 71:375–510.

Smith, A.C. 1932. The American species of Thibaudieae. Contr. U.S. Natl. Herb. 28:311–547.

Smith, A.C. 1933. Studies of South American plants, III. New Ericaceae and Vacciniaceae. Bull. Torrey Bot. Club 60:99–121. [Satyria cerander (Dunal) A.C.Sm., p. 120. Doi: 10.2307/2480601]

Smith, A.C. 1935. Studies of South American plants, IV. New Monimiaceae, Trigoniaceae, and Vacciniaceae. Phytologia 1:126–132.

Smith, A.C. 1941. Vacciniaceae. In: Woodson, R.E., Jr. & R.W. Schery, eds. Contributions toward a Flora of Panama. V. Ann. Missouri Bot. Gard. 28:438–452.

Smith, A.C. 1946. Studies of South American plants, XI. Noteworthy species of Hippocrateaceae and Vacciniaceae. J. Arnold Arbor. 27:86–120.

Smith, A.C. 1952. Plants collected in Ecuador by W.H. Camp—Vacciniaceae. Mem. New York Bot. Gard. 8:41–85.

Soukup, J. 1972. Las Cletráceas, Ericáceas, Teofrastáceas, Mirsináceas, Primuláceas y Plumbagináceas del Perú, sus géneros y lista de especies. Biota 9. [Ericaceae, pp. 224–239]

Stafleu, F.A. 1967. Taxonomic literature. Regnum Veg. 52. Utrecht.

Stafleu, F.A. & R.S. Cowan. 1976–1988. Taxonomic literature: a selective guide to botanical publications and collections with dates, commentaries and types. Ed. 2, 6 volumes. Bohn, Scheltema & Holkema, Utrecht. Https://doi.org/10.5962/bhl.title.48631 [Vol. 2 Authors H–Le (1979); 3 Authors Lh–O (1981); 4 Authors P–Sak (1983)]

Stauffer, F.W., J. Stauffer & L.J. Dorr. 2012. The Bonpland and Humboldt monocotyledon collections based on the study of the specimens gathered during their Venezuelan exploration. Candollea 67:75–130.

Steele, A.R. 1964. Flowers for the king: the expedition of Ruiz and Pavón and the Flora of Peru. Duke University Press, Durham, NC.

Steinberg, C.H. 1977. The collectors and collections in the Herbarium Webb. Webbia 32:1–49.

Steyermark, J.A. 1981. Erroneous citations of Venezuelan localities. Taxon 30(4):816–817.

Steyermark, J.A., P.E. Berry, & B. Holst. 1995. Flora of the Venezuelan Guyana, vol. 1: Introduction. Missouri Botanical Garden Press. St. Louis, Missouri.

Stuessy, T.F. 2009. Plant taxonomy: the systematic evaluation of comparative data. Ed. 2. Columbia University Press, New York.

Turland, N.J., J.H. Wiersema, F.R. Barrie, W. Greuter, D.L. Hawksworth, P.S. Herendeen, S. Knapp, W.-H. Kusber, D.-Z. Li, K. Marhold, T.W. May, J. Mcneill, A.M. Monro, J. Prado, M.J. Price, & G.F. Smith, eds. 2018. International code of nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Veg. 159. Glashütten: Koeltz Botanical Books. Https://doi.org/10.12705/Code.2018

Urban, I., ed. 1902. Symbolae antillanae seu fundamenta florae Indiae Occidentalis. [“II. Notae biographicae peregrinatorum Indiae occidentalis botanicorum,” vol. 3(fasc. 1):14–160.] Fratres Borntraeger, Leipzig. Paul Klincksieck, Paris. Williams & Norgate, London. 1898–1928. [see pp. 49–50, 76–78, and 120–121 for notes on Funck, Linden, & Schlim, respectively]

Van Dam, J.A.C. 2002. The Guyanan plant collections of Robert and Richard Schomburgk. In: M.J. Jansen-Jacobs, ed. Flora of the Guianas. Supplementary Series Fascicle 3. Royal Botanic Gardens, Kew. Richmond, Surrey, U.K.

Vegter, I.H. 1976. Index Herbariorum: Part II(4), Collectors, M. Regnum Veg. 93. Bohn, Scheltema & Holkema, Utrecht.

Vegter, I.H. 1986. Index Herbariorum: Part II(6), Collectors, S. Regnum Veg. 114. Bohn, Scheltema & Holkema, Utrecht.

Vesque, J. 1885. Caractères des principales familles Gamopétales: Tirés de L’Anatomie de la feuille. Ann. Sci. Nat. Bot., ser. 7, t. 1:183–360 [Vacciniaceae, pp. 240–244, Psammisia planchoniana pp. 240, 242]

Walpers, G.G. 1848–68. Annales botanices systematicae. Sumtibus F. Hofmeister, Lipsiae. [Ericaceae s.l., vol. 2:1077–1125 (1851)]

Werner, F.A. & G. Mendieta-Leiva. 2014. Ericaceae of the “Estacio?n Cienti?fica San Francisco.” Rapid Color Guide # 630 versio?n 1. 7/2014. The Field Museum.

Wilbur. R.L. & J.L. Luteyn. 1978. Family 149 Ericaceae. In: R.E. Woodson et collab., eds. Flora of Panama, Part VIII. Ann. Missouri Bot. Gard. 65:27–144.

Wurdack, J.J. 1964. In Mathew's country. J. New York Bot. Gard. 14:7–10.



How to Cite

Luteyn, J. L. (2019). Typification in and contributions to a revision of Psammisia (Ericaceae: Vaccinieae). Journal of the Botanical Research Institute of Texas, 13(2), 397–449. https://doi.org/10.17348/jbrit.v13.i2.796